Abstract ........................................................ 4
Introduction .................................................... 5
Abbreviations ................................................... 6
Platybasic and Tropibasic Braincases ............................ 8
"Cobelodus" .................................................... 18
Material Examined .......................................... 18
General Remarks ............................................ 18
General Morphology ......................................... 19
Ethmoid Region ............................................. 29
Orbit and Interorbital Septum .............................. 30
Supraorbital Shelf ......................................... 31
Suborbital Shelf ........................................... 31
Optic Nerve, Optic Pedicel, Oculomotor Nerve, and
Pituitary Vein ......................................... 31
Trochlear Nerve ............................................ 32
External Rectus Muscle ..................................... 32
Trigeminal, Profundal, and Anterodorsal Lateral Line
Nerves ................................................. 33
Facial and Abducent Nerves ................................. 35
Postorbital Arcade ......................................... 36
Arterial Supply to the Head ................................ 37
Otic Region ................................................ 41
Occipital Region ........................................... 42
Otico-occipital Fissure .................................... 45
Spino-occipital Nerves ..................................... 45
Cranial Endocast ........................................... 48
Diencephalon and Hypophyseal Chamber ....................... 56
Labyrinth Endocast ......................................... 58
Other Symmoriiform Remains from Arkansas ....................... 59
Material Examined .......................................... 59
General Remarks ............................................ 59
Description ................................................ 60
Virgilian Symmoriiform Braincase ............................... 62
Material Examined .......................................... 62
Description ................................................ 62
Cobelodus aculeatus (Cope 1894) ................................ 63
Material Examined .......................................... 63
General Remarks ............................................ 63
Concretion Specimen ........................................ 64
X-rays ..................................................... 69
Compression Specimens ...................................... 71
Symmorium reniforme Cope, 1893 ................................. 77
Material Examined .......................................... 77
General Remarks ............................................ 77
Description ................................................ 79
Stethacanthulus meccaensis (Williams, 1985) .................... 80
Material Examined .......................................... 80
General Remarks ............................................ 80
Description ................................................ 81
Late Pennsylvanian Symmoriiforms from New Mexico ............... 81
General Remarks ............................................ 81
Cranial Morphology ......................................... 81
Denaea fournieri Pruvost (in Fournier and Pruvost, 1922) ....... 83
General Remarks ............................................ 83
Akmonistion zangerli Coates and Sequeira, 2001a ................ 84
General Remarks ............................................ 84
Cranial Morphology ......................................... 84
"Stethacanthus" ................................................ 85
General Remarks ............................................ 85
Cranial Morphology ......................................... 85
Dwykaselachus oosthuiseni Oelofsen, 1986 ....................... 89
General Remarks ............................................ 89
Cranial Morphology ......................................... 89
Falcatus falcatus (St. John and Worthen, 1883) ................. 90
Material Examined .......................................... 90
General Remarks ............................................ 90
Description ................................................ 94
Damocles serratus Lund, 1986 ................................... 96
Material Examined .......................................... 96
General Remarks ............................................ 96
Description ................................................ 96
Cladoselache Dean, 1894 ........................................ 97
General Remarks ............................................ 97
Cranial Morphology ......................................... 97
Discussion .................................................... 102
Tropibasia in Symmoriiforms ............................... 102
Scleral Ossicles .......................................... 105
A Reconstruction of the Jaws in "Cobelodus" ............... 106
The Anterior Palatoquadrate Articulation .................. 109
The Postorbital Articulation .............................. 110
Hyomandibular Involvement in Jaw Suspension ............... 112
Conclusions ................................................... 114
Acknowledgments ............................................... 116
References .................................................... 116
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