LIST OF COLOUR PAINTINGS ..................................... viii
PREFACE ........................................................ ix
Author's Note ............................................... ix
Acknowledgements ............................................. x
Rowans - looking to the future ............................. xii
1 INTRODUCTION ................................................. 1
Brief account of the Rosaceae ................................ 1
Sorbus in cultivation ........................................ 3
Chromosome numbers in the Rosaceae ........................... 4
The subfamily Maloideae ...................................... 5
The genus Sorbus (the pinnate-leaved rowans) ................. 6
Sexual and asexual reproduction in Sorbus .................... 6
Sexual species ............................................ 6
Asexual species: apomixis ................................. 7
The naming of apomicts .................................... 8
How microspecies arise - Sorbus × Aria .................... 9
Origin of the Sino-Himalayan apomictic microspecies of
Sorbus ................................................... 11
Evolution of Sorbus ...................................... 11
2 PHYLOGEOGRAPHY .............................................. 15
Fossil evidence ............................................. 15
Distribution of taxa within Sorbus .......................... 16
Comparisons with the fossil history of other genera with
which Sorbus species often grow ............................. 18
Cenozoic Tertiary fossil record of associated genera ..... 19
The Eocene period ..................................... 19
Oligocene and Miocene ................................. 22
Pliocene and Pleistocene .............................. 24
Evolution and dispersal of Sorbus aucuparia throughout
northern Eurasia ......................................... 25
The value of traditional taxonomy to molecular taxonomy .. 27
3 ECOLOGY ..................................................... 28
4 CONSERVATION ................................................ 29
5 CULTIVATION ................................................. 31
Introduction ................................................ 31
Temperature ................................................. 31
Moisture .................................................... 32
pH and nutrients ............................................ 33
Pests and diseases .......................................... 35
Mammals .................................................. 35
Insects .................................................. 35
Bird predation of fruit .................................. 36
Fungi .................................................... 37
Herbicides .................................................. 39
Propagation ................................................. 40
Grafting ................................................. 40
Cuttings ................................................. 40
Seed ..................................................... 41
Seed extraction and cleaning .......................... 43
Germination ........................................... 43
Choosing a suitable rowan for your garden ................... 46
6 TAXONOMIC TREATMENT ......................................... 49
Infrageneric classification ................................. 49
Subgenus Sorbus .......................................... 50
Subgenus Albocarmesinae .................................. 50
Morphology .................................................. 51
Habit .................................................... 51
Shoots and twigs ......................................... 51
Bud ...................................................... 52
Leaf ..................................................... 53
Inflorescence ............................................ 55
Flower ................................................... 55
Fruit .................................................... 56
Summary of classification ................................... 61
Subgeneric and sectional classification .................. 61
Arrangement of the species ............................... 64
Key to the species of Sorbus ................................ 65
The Genus Sorbus ............................................ 73
Subgenus Sorbus (Species 1-17) ........................... 73
Section Sorbus (Species 1-3) .......................... 73
Section Commixtae (Species 4-9) ....................... 85
Section Wilsoniae (Species 10-11) ..................... 94
Section Sambucifoliae (Species 12) .................... 96
Section Graciles (Species 13) ......................... 98
Section Tianshanicae (Species 14-17) .................. 99
Subgenus Albocarmesinae (Species 18-72) ................. 104
Section Discolores (Species 18-30) ................... 104
Section Insignes (Species 31-33) ..................... 130
Section Hypoglaucae (Species 34-39) .................. 134
Section Multijugae (Species 40-69) ................... 140
Section Reductae (Species 70-72) ..................... 209
Hybrids ................................................. 216
APPENDIX: Latin descriptions of New Taxa ...................... 231
GLOSSARY and ABBREVIATIONS .................................... 239
BIBLIOGRAPHY .................................................. 245
INDEX OF SCIENTIFIC NAMES ..................................... 251
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