INTRODUCTION
PART I PALYNOSTRATIGRAPHY OF THE SOUTH PORTUGUESE ZONE AND
TECTONO-STRATIGRAPHIC IMPLICATIONS
1 Stratigraphic Setting ....................................... 15
1.1 Pulo do Lobo Domain .................................... 15
1.2 Iberian Pyrite Belt (IPB) .............................. 16
1.3 Baixo Alentejo Flysch Group (BAFG) ..................... 16
1.4 The Southwest Portugal Domain (SWP) .................... 17
Carrapateira Group .......................................... 17
2 Previous Palynostratigraphic Studies in the South
Portuguese Zone - Historical Review ......................... 19
3 Methods ..................................................... 20
3.1 Samples Preparation .................................... 20
3.2 Miospore Zonation Scheme ............................... 21
4 Palynostratigraphic framework of the South Portuguese Zone .. 23
4.1 Pulo do Lobo Antiform .................................. 23
Pulo do Lobo and Atalaia Formations ......................... 23
Ribeira de Limas and Gafo Formations ........................ 23
Santa Iria, Horta da Torre and Represa Formations ........... 23
4.2 Iberian Pyrite Belt .................................... 27
North Branch ........................................... 27
i Chaparral region ................................... 27
ii Alberrtôa and Serra Branca regions ................. 27
iii Azenhas Section, Guadiana River (Mértola) .......... 33
iv São Domingos mine boreholes and open pit section ... 35
South Branch ........................................... 38
i Pomarão Anticline .................................. 38
ii Neves Corvo mine region ............................ 38
iii Aljustrel mine region .............................. 42
iv Lousal/Azinheira de Barros/Mina da Caveira
regions ............................................ 43
v São Francisco da Serra region ...................... 45
vi Cereal Anticline region ............................ 45
4.3 The Baixo Alentejo Flysch Group ........................ 51
Mértola Formation ...................................... 51
Mira Formation ......................................... 51
Brejeira Formation ..................................... 51
4.4 Southwest Portugal (SWP) Domain, Bordeira and Aljezur
Anticlines ............................................. 52
Tercenas Formation ..................................... 52
Bordalete Formation .................................... 52
Murração Formation ..................................... 54
Quebradas Formation .................................... 54
5 Reworked Palynomorphs ....................................... 54
5.1 Reworked Palynomorphs in the Devonian units of SPZ ..... 57
5.2 Reworked palynomorphs in the Carboniferous units of
SPZ .................................................... 57
6 Correlation with Europe ..................................... 58
7 Synopsis of the South Portuguese Zone palynostratigraphy .... 59
8 Implications on the SPZ tectonostratigraphic and
geodynamic interpretation ................................... 60
PART 2 SYSTEMATIC PALYNOLOGY
1 Miospores ................................................... 65
Anteturma SPORITES H. Potonié 1893 .......................... 65
Turma TRILETES (Reinsch) Dettmann 1963 ...................... 65
Suprasubturma ACAMERATITRILETES Neves & Owens 1966 .......... 65
Subturma AZONOTRILETES (Luber) Dettmann, 1963 ............... 65
Infraturma LAEVIGATI Bennie & Kidston emend R. Potonié
1956 ........................................................ 65
Genus CALAMOSPORA Schopf, Wilson & Bentall 1944 .......... 65
Genus LEIOTRILETES Naumova ex Ishchenko 1952 ............. 65
Genus PLICATISPORA Higgs, Clayton & Keegan, 1988 ......... 66
Genus PUNCTATISPORITES Ibrahim emend R. Potonié & Kremp,
1954 ..................................................... 66
Infraturma RETUSOTRILETI Streel in Becker, Bless, Street &
Thorez, 1974 ................................................ 67
Genus RETUSOTRILETES Naumova emend Streel, 1964 .......... 67
Infraturma APICULATI Bennie & Kidston emend R. Potonié
1956 ..................................................... 69
Genus ACANTHOTRILETES Naumova emend Potonié &
Kremp 1954 ............................................... 69
Genus ANAPICULATISPORITES (Potonié & Kremp) emend
Smith & Butterworth 1967 ................................. 70
Genus ANAPLANISPORITES Jansonius 1962 .................... 70
Genus APICULIKETUSISPORA Streel emend Streel 1967 ........ 71
Genus CADIOSPORA (Kosanke) Venkatachala & Bharadwaj
1964 ..................................................... 71
Genus CAMPTOTRILETES Naumova emend Potonié & Kremp 1954 .. 71
Genus CRASSISPORA Bharadwaj emend Sullivan 1964 .......... 72
Genus GRANULATISPORITES Ibrahim emend Potonié & Kremp
1954 ..................................................... 73
Genus LOPHOTRILETES (Naumova) Potonié & Kremp 1954 ....... 73
Genus NEORAISTRICKIA Potonié 1956 ........................ 74
Genus PUSTULATISPORITES R. Potonié & Kremp emend Imgrund
1960 ..................................................... 74
Genus RAISTRICKIA Schopf, Wilson & Bentall emend R.
Potonié & Kremp 1954 ..................................... 75
Genus SCHOPFITES Kosanke 1950 ............................ 76
Genus UMBONATISPORITES Hibbert & Lacey 1969 .............. 77
Genus VERRUCIRETUSISPORA Owens 1971 ...................... 77
Genus VERRUCOSISPORITES Ibrahim emend Smith &
Butterworth, 1967 ........................................ 77
Genus WALTZISPORA Staplin 1960 ........................... 79
Infraturma MURORNATI R Potonié & Krempl954 ................. 80
Genus ACINOSPORITES Richardson 1965 ...................... 80
Genus CONVOLUTISPORA Hoffmeister, Staplin & Malloy,
1955 ..................................................... 80
Genus DICTYOTRILETES Naumova emend Smith & Butherworth
1967 ..................................................... 81
Genus EMPHANISPORITES McGregor, 1961 ..................... 82
Genus MICRORETICULATISPORITES Knox emend Potonié &
Kremp 1954 ............................................... 83
Subturma ZONOTRILETES Waltz, 1935 ........................... 83
Infraturma AURICULATI Smith & Butterworth, 1967 ............. 83
Genus TRI PARTITES (Schemel) Potonié & Kremp 1954 ........... 84
Genus TRIQUITRITES (Wilson & Сое) Potonié & Kremp 1954 ... 84
Infraturma TRICRASSATI Dettmann 1963 ........................ 84
Genus DIATOMOZONOTRILETES (Naumova) Playford 1963 ........ 84
Infraturma CINGULATI Potonié & Klaus emend Dettmann, 1963 ... 85
Genus ANEUROSPORA Streel, 1964 ........................... 85
Genus BELLISPORES (Artuz) Sullivan 1964 .................. 86
Genus KNOXISPORITES R. Potonié & Kremp emend Neves 1961 .. 86
Genus ROTASPORA Schemel emend. Smith & Butterworth 1967 .. 88
Genus RETICULATISPORITES (Ibrahim) Neves 1964 ............ 89
Genus SAVITRISPORITES Bharadwaj 1955 ..................... 90
Genus SECARISPORITES Neves 1961 .......................... 91
Genus TUMULISPORA Staplin & Jansonius, 1964 .............. 91
Suprasubturma LAMINATITRILETES Smith & Butterworth, 1967 .... 91
Subturma ZONOLAMINATITRILETES Smith & Butterworth, 1967 ..... 91
Infraturma PATIN ATI Butterworth & Williams, 1958 ........... 91
Genus ARCHAEOZONOTRILETES (Naumova) Allen 1965 ........... 91
Genus CHELINOSPORA Allen 1965 ............................ 92
Genus CYRTOSPORA Winslow 1962 ............................ 92
Genus CYMBOSPORITES Allen 1962 ........................... 93
Infraturma CINGULICAVATI Smith & Butterworth, 1967 .......... 93
Genus CAMAROZONOTRILETES Naumova 1953 .................... 93
Genus CINGULIZONATES (Dybova & Jachowicz) Butterworth,
Jansonius, Smith & Staplin 1964 .......................... 94
Genus CIRRATRIRADITES Wilson & Сое 1940 .................. 94
Genus CRISTATISPORITES Potonié & Kremp emend
Butterworth, Jansonius, Smith & Staplin 1964 ............. 94
Genus DENSOSPORITES Berry emend Butterworth, Jansonius,
Smith & Staplin 1964 ..................................... 95
Genus INDOTRIRADIATES Tiwari emend. Foster, 1979 ......... 96
Genus KRAEUSELISPORITES Leschik emend Jansonius 1962 ..... 97
Genus LYCOSPORA Schopf, Wilson & Bentall emend R.
Potonié & Kremp, 1954 .................................... 98
Genus RADIIZONATES Staplin & Jansonius 1964 .............. 98
Genus VALLATISPORITES Hacquebard 1957 emend Sullivan
1964 ..................................................... 98
Suprasubturma PSEUDOSACCITRILETES Richardson, 1965 ...... 100
Infraturma MONOPSEUDOSACCITI Smith & Butterworth, 1967 .. 100
Genus ANCYROSPORA Richardson emend. Richardson 1962 ..... 100
Genus AURORASPORA Hoffmeister, Staplin & Malloy 1955 .... 100
Genus DIDUCITES Van Veen 1981 ........................... 101
Genus CRISTICAVATISPORA González, Moreno & Playford
2005 .................................................... 102
Genus ENDOCULEOSPORA Staplin emend Turnau 1975 .......... 103
Genus ENDOSPORITES Wilson & Сое ex Schopf, Wilson &
Bentall 1944 ............................................ 103
Genus EPIGRUSPORA González, Moreno & Playford 2005 ...... 104
Genus GEMINOSPORA Balme 1962 ............................ 104
Genus GRANDISPORA Hoffmeister, Staplin & Malloy emend.
McGregor 1973 ........................................... 105
Genus RETISPORA Staplin 1960 ............................ 107
Genus RUGOSPORA Neves & Owens 1966 ...................... 108
Genus SPELAEOTRILETES Neves & Owens 1966 ................ 109
Genus TEICHERTOSPORA Balme 1988 ......................... 110
Infraturma MEMBRANATI Neves 1961 ........................... 110
Genus PROPISPORITES Neves 1958 .......................... 110
Turma MONOLETES Ibrahim 1933 ............................... 111
Subturma AZONOMONOLETES Luber 1935 ......................... 111
Infraturma PSILAMONOLETI van der Hammen .................... 111
Genus LATOSPORITES Potonié & Kremp 1954 ................. 111
Infraturma SCULPTATOMONOLETI Dybová & Jachowicz 1957 ....... 111
Genus ARATRISPORITES Leschik 1955 emend Playford &
Dettman 1965 ............................................ 111
Genus THYMOSPORA Wilson & Venkatachala 1963 ............. 111
Genus TORISPORA (Balme) Doubinger & Horst 1961 .......... 112
Anteturma POLLENITES Potonié 1931 .......................... 112
Turma Saccites Erdtman 1947 ................................ 112
Subturma MONOSACCITES Potonié & Kremp 1954 ................. 112
Infraturma ARADITES Schopf, Wilson & Bentall 1944 .......... 112
Genus FLORINITES Schopf, Wilson & Bentall 1944 .......... 112
INSERTAESEDIS .............................................. 112
Genus COLATISPORITES Williams em Neves et al. 1973 ...... 112
2 Prasinophytes and Acritarchs ............................... 113
Division CHLOROPHYTA Pascher, 1914 ......................... 113
Class PRASINOPHYCEAE Christensen, 1962 ..................... 113
Family LEIOSPHAERIDIACEAE Timofeev, 1956 nom. corr.
Madler, 1963 ............................................... 113
Genus LEIOSPHAERIDIA Eisenack, 1958 ..................... 113
Genus TASMANITES Newton, 1875 ........................... 113
Family CYMATIOSPHAERACEAE Mädler, 1963 ..................... 113
Genus CYMATIOSPHAERA O. Wetzel ex Deflandre, 1954 ....... 113
Genus DICTYOTIDIUM Eisenack emend. Staplin, 1961 ........ 114
Genus DUVERNAYSPHAERA (Staplin) Deunff 1964 ............. 114
Family PTEROSPERMELLACEAE Eisenack, 1972 ................... 115
Genus MARANHITES Brito, 1965 ............................ 115
Genus PTEROS Per MELLA Eisenack 1972 .................... 117
Group ACRITARCHA Evitt, 1963 ............................... 118
Genus CHOMOTRILETES Naumova ex Naumova, 1953 ............ 118
Genus CRASSIANGULINA Jardiné, Combaz, Magloire,
Peninguel & Vachey, 1972 ................................ 118
Genus CRATERISPHAERIDIUM Deunff, 1981 ................... 119
Genus DAILLYDIUM Stockmans & Willière, 1969 ............. 119
Genus DELTOTOSOMA Playford, 1981 ........................ 120
Genus DUPLICIRADIATUM González, Moreno & Playford,
2005 .................................................... 120
Genus GORGONISPHAERIDIUM Staplin, Jansonius & Pocock
1965 .................................................... 121
Genus HOROLOGINELLA Coockson & Eisenack 1962 ............ 122
Genus MICRHYSTRIDIUM Deflandre, 1937 .................... 122
Genus NAVIFUSA Combaz, Lange & Pansart 1967 ............. 122
Genus PALACANTHUS Wicander emend. Sarjeant &
Stancliffe, 1994 ........................................ 123
Genus STELUNIUM Jardiné, Combaz, Magloire, Peniguel &
Vachey, 1972 ............................................ 124
Genus UMBELLASPHAERIDIUM Jardiné, Combaz, Magloire,
Peniguel & Vachey, 1972 ................................. 124
Genus UNELLIUM Rauscher, 1969 ........................... 125
Genus VERYHACHIUM Deunff ex Downie 1959 emend.
Sarjeant & Stancliffe, 1994 ............................. 126
Genus WINWALOEUSIA Deunff, 1977 ......................... 127
REFERENCES .................................................... 129
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