Introduction .................................................... 7
Acknowledgements ................................................ 8
Previous research ............................................... 9
The Pliensbachian Stage in the Bakony Mountains ................ 11
Stratigraphy and palaeogeography ............................ 11
Sedimentary environments .................................... 13
Lithologies .............................................. 13
Interpretation ........................................... 17
Sections and localities ..................................... 18
Brachiopod fauna ............................................ 37
Systematic palaeontology ....................................... 41
Methods ..................................................... 41
Systematic descriptions ..................................... 42
Order Rhynchonellida Kuhn, 1949 .......................... 42
Superfamily Pugnacoidea Rzhonsnitskaia, 1956 .......... 42
Family Basiliolidae Cooper, 1959 ................... 42
Subfamily Basiliolinae Cooper, 1959 ............. 42
Genus Apringia de Gregorio, 1886 ............. 42
Apringia piccininii (Zittel, 1869) ........ 43
Apringia barnabasi n. sp .................. 45
Apringia paolii (Canavari, 1880) .......... 46
Apringia diptycha (Böse, 1898) ............ 50
Apringia deltoidea (Canavari, 1880) ....... 53
Genus Serratapringia n. gen .................. 53
Serratapringia fraudatrix (Böse, 1898) .... 54
Serratapringia? cf. suetii (Haas, 1884) ... 55
Genus Megapringia n. gen ..................... 56
Megapringia stoppanii (Parona, 1880) ...... 57
Megapringia altesinuata (Böse, 1898) ...... 59
Megapringia? atlaeformis (Böse, 1898) ..... 61
Subfamily Pamirorhynchiinae Ovcharenko, 1983 .... 63
Genus Jakubirhynchia Tomašovỳch, 2006 ........ 63
Jakubirhynchia? cf. fascicostata (Uhug,
1880) ..................................... 63
Jakubirhynchia? aff. laevicosta
(Geyer, 1889) ............................. 64
Superfamily Rhynchotetradoidea Licharew, 1956 ......... 65
Family Prionorhynchiidae Manceñido & Owen, 2002 .... 65
Genus Prionorhynchia Buckman, 1918 ........... 65
Prionorhynchiapolyptycha (Oppel, 1861) .... 65
Prionorhynchia aff. greppini (Oppel,
1861) ..................................... 66
Prionorhynchia? hagaviensis (Böse,
1898) ..................................... 67
Prionorhynchia? flabellum (Gemmellaro,
1874) ..................................... 68
Prionorhynchia? catharinae n. sp .......... 70
Genus Lokutella Vörös, 1983 .................. 71
Lokutella liasina (Principi, 1910) ........ 71
Lokutella palmaeformis (Haas, 1912) ....... 73
Lokutella kondai Vörös, 1983 .............. 75
Superfamily Wellerelloidea Licharew, 1956 ............. 76
Family Wellerellidae Licharew, 1956 ................ 76
Subfamily Cirpinae Ager, 1965 ................... 76
Genus Cirpa de Gregorio, 1930 ................ 76
Cirpa? subcostellata (Gemmellaro,
1878) ..................................... 76
Cirpa? cf. subfurcillata (Böse, 1898) ..... 78
Genus Calcirhynchia Buckman, 1918 ............ 78
Calcirhynchia? hungarica (Böckh, 1874) .... 78
Superfamily Rhynchonelloidea D'Orbigny, 1847 .......... 80
Family Rhynchonellidae D'Orbigny, 1847 ............. 80
Subfamily Rhynchonellinae D'Orbigny, 1847 ....... 80
Genus Homoeorhynchia Buckman, 1918 ........... 80
Homoeorhynchia cf. acuta (J. Sowerby,
1816) ..................................... 80
Homoeorhynchia? lubrica (Uhlig, 1880) ..... 81
Homoeorhynchia? ptinoides (Di Stefano,
1891) ..................................... 83
Subfamily Piarorhynchiinae Shi & Grant, 1993 .... 84
Genus Cuneirhynchia Buckman, 1918 ............ 84
Cuneirhynchia cf. rastuensis Benigm,
1978 ...................................... 84
Superfamily Norelloidea Ager, 1959 .................... 84
Family Norellidae Ager, 1959 ....................... 84
Subfamily Norellinae Ager, 1959 ................. 84
Genus Pisirhynchia Buckman, 1918 ............. 84
Pisirhynchia pisoides (Zittel, 1869) ...... 84
Pisirhynchia retroplicata (Zittel,
1869) ..................................... 86
Genus Fenyveskutella n. gen .................. 87
Fenyveskutella vighi n. sp ................ 88
Fenyveskutella pseudouhligi n. sp ......... 90
Fenyveskutella theresiae n. sp ............ 91
Fenyveskutella aff. vighi n. sp ........... 92
Genus Kericserella Vörös, 1983 ............... 92
Kericserella inversaeformis (Schlosser,
1900) ..................................... 93
Subfamily Laevirhynchiinae Dagys, 1974 ................ 94
Genus Nannirhynchia Buckman, 1918 ............ 94
Nannirhynchia reynesi (Gemmellaro,
1874) ..................................... 95
Nannirhynchia gemmellaroi (Parona,
1880) ..................................... 96
Nannirhynchia sinuata (Haas, 1912) ........ 98
Genus Paronarhynchia n. gen .................. 99
Paronarhynchia bulga (Parona, 1893) ....... 99
Paronarhynchia estherae n. sp ............ 101
Paronarhynchia? cf. verrii (Parona,
1883) .................................... 102
Superfamily Hemithyridoidea Rzhonsnitskaia, 1956 ..... 103
Family Tetrarhynchiidae Ager, 1959 ................ 103
Subfamily Gibbirhynchiinae Manceńido &
Owen, 2002 ..................................... 103
Genus Gibbirhynchia Buckman, 1918 ........... 103
Gibbirhynchia cf. curviceps (Quenstedt,
1858) .................................... 103
Gibbirhynchia? orsinii (Gemmellaro,
1874) .................................... 104
Gibbirhynchia? cf. urkutica (Böckh,
1874) .................................... 106
Order Athyridida Boucot, Johnson & Staton, 1964 ......... 107
Suborder Koninckinidina Harper, 1993 ................. 107
Superfamily Koninckinoidea Davidson, 1853 ......... 107
Family Koninckinidae Davidson, 1853 ............ 107
Genus Amphiclinodonta Bittner, 1888 ......... 107
Amphiclinodonta Hasina Bittner, 1888 ..... 107
Genus Koninckodonta Bittner, 1894 ........... 108
Koninckodonta cf. waehneri (Bittner,
1894) .................................... 108
Koninckodontafuggeri Bittner, 1894 ....... 109
Koninckodonta eberhardi Bittner, 1888 .... 110
Order Spiriferinida Ivanova, 1972 ....................... 111
Suborder Spiriferinidina Ivanova, 1972 ............... 111
Superfamily Pennospiriferinoidea Dagys, 1972 ....... 111
Family Lepismatinidae Xu & Liu, 1983 ............. 111
Subfamily Dispiriferininae Carter, 1994 ........ 111
Genus Dispiriferina Siblнk, 1965 ............ 111
Dispiriferina cf. segregata
(Di Stefano, 1887) ....................... 111
Superfamily Spiriferinoidea Davidson, 1884 ......... 112
Family Spiriferinidae Davidson, 1884 ............. 112
Subfamily Spiriferininae Davidson, 1884 ........ 112
Genus Liospiriferina Rousselle, 1977 ........ 112
Liospiriferina alpina (Oppel, 1861) ...... 112
Liospiriferina cf. brevirostris (Oppel,
1861) .................................... 114
Liospiriferina obtusa (Oppel, 1861) ...... 115
Liospiriferina sicula (Gemmellaro,
1874) .................................... 117
Liospiriferina gryphoidea (Uhlig,
1880) .................................... 119
Liospiriferina apenninica (Canavari,
1880) .................................... 121
Liospiriferina aff. handeli (Di
Stefano, 1887) ........................... 121
Liospiriferina cf. semicircularis (Böse,
1898) .................................... 122
Liospiriferina cf. globosa (Böse,
1898) .................................... 124
Liospiriferina cf. obovata (Principi,
1910) .................................... 124
Subfamily Paralaballinae Carter, 1994 .......... 125
Genus Cisnerospira Manceñido, 2004 .......... 125
Cisnerospira darwini (Gemmellaro,
1878) .................................... 125
Cisnerospira meneghiniana (Canavari,
1880) .................................... 126
Order Terebratulida Waagen, 1883 ........................ 127
Suborder Terebratulidina Waagen, 1883 ................ 127
Superfamily Uncertain ............................. 127
Family Orthotomidae Muir-Wood, 1936 ............ 127
Genus Orthotoma Quenstedt, 1869 ............. 127
Orthotoma apenninica (Canavari, 1883) .... 127
Orthotoma aff. apenninica (Canavari,
1883) .................................... 129
Superfamily Loboidothyridoidea Makridin, 1964 ..... 129
Family Plectoconchidae Dagys, 1974 ............. 129
Genus Lychnothyris Vörös, 1983 .............. 129
Lychnothyris rotzoana (Schauroth,
1865) .................................... 130
Genus Merophricus Cooper, 1983 .............. 131
Merophricus cf. mediterraneus
(Canavari, 1883) ......................... 132
Merophricus aff. moreti (Dubar, 1942) .... 133
Family Hesperithyrididae Cooper, 1983 .......... 133
Genus Hesperithyris Dubar, 1942 ............. 133
Hesperithyris renierii (Catullo, 1827) ... 134
Hesperithyris cf. costata (Dubar,
1942) .................................... 137
Family Lobothyrididae Makridin, 1964 ........... 137
Subfamily Lobothyridinae Makridin, 1964 ....... 137
Genus Lobothyris Buckman, 1918 .............. 137
Lobothyrispunctata (J. Sowerby, 1813) .... 137
Genus Rhapidothyris Tuluweit, 1965 .......... 139
Rhapidothyris cf. ovimontana (Böse,
1898) .................................... 140
Rhapidothyris delorenzoi (BÖSE, 1900) .... 142
Rhapidothyris beyrichi (Oppel, 1861) ..... 142
Rhapidothyris lokutica n. sp ............. 145
Family Uncertain ............................... 148
Genus Viallithyris Vörös, 1978 .............. 148
Viallithyris gozzanensis (Parona,
1880) .................................... 148
Genus Eplenyithyris n. gen .................. 153
Eplenyithyris cerasulum (Zittel, 1869) ... 154
Superfamily Dyscolioidea Fischer & Oehlert,
1891 .............................................. 157
Family Pygopidae Muir-Wood, 1965 ................. 157
Subfamily Triangopinae Manceńido, 1993 ......... 157
Genus Securithyris Vörös, 1983 .............. 157
Securithyris adnethensis (Suess, 1855) ... 158
Securithyrisfilosa (Canavari, 1880) ...... 165
Securithyris paronai (Canavari, 1880) .... 167
Family Nucleatidae Schuchert, 1929 ............. 168
Genus Linguithyris Buckman, 1918 ............ 168
Linguithyris aspasia (Zittel, 1869) ...... 169
Linguithyris cf. linguata (Böckh,
1874) .................................... 177
Genus Buckmanithyris Tchorszhevsky, 1990 .... 178
Buckmanithyris cornicolana (Canavari,
1881) .................................... 178
Family Uncertain ............................... 179
Genus Papodina Vörös, 1983 .................. 179
Papodina bittneri (Geyer, 1889) .......... 179
Suborder Terebratellidina Muir-Wood, 1955 ............ 183
Superfamily Zeillerioidea Allan, 1940 .............. 183
Family Zeilleriidae Allan, 1940 .................. 183
Subfamily Zeilleriinae Allan, 1940 ............. 183
Genus Zeilleria Bayle, 1878 ................. 183
Zeilleria mutabilis (Oppel, 1861) ........ 183
Zeilleria alpina (Geyer, 1889) ........... 186
Zeilleria bicolor (Böse, 1898) ........... 188
Zeilleria? aquilina (Franceschi, 1921) ... 189
Genus Antiptychina Zittel, 1880 ............. 191
Antiptychina? rothpletzi (Di Stefano,
1891) .................................... 191
Genus Aulacothyris Douvillé, 1879 ........... 193
Aulacothyris? cf. fuggeri (Böse, 1898) ... 193
Aulacothyris? ballinensis (Haas, 1912) ... 193
Genus Bakonyithyris Vörös, 1983 ............. 196
Bakonyithyris pedemontana (Parona,
1893) .................................... 196
Bakonyithyris apenninica (Zittel,
1869) .................................... 198
Bakonyithyris avicula (Uhlig, 1880) ...... 201
Bakonyithyris meneghinii (Parona,
1880) .................................... 203
Bakonyithyris ovimontana (Böse, 1898) .... 204
Bakonyithyris? aff. gastaldii (Parona,
1880) .................................... 207
Genus Securina Vörös, 1983 .................. 208
Securina hierlatzica (Oppel, 1861) ....... 208
Appendix ...................................................... 211
References .................................................... 225
Plates ........................................................ 237
Index ......................................................... 297
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